Family: Rhabdoviridae

Genera not assigned to a subfamily

Within the Rhabdoviridae, seven genera do not fall within any of the three existing subfamilies. All viruses assigned to these genera have been detected by high-throughput sequencing of metagenomes of invertebrates.

Genus Alphacrustrhavirus. The viruses in this genus have been detected in marine crustaceans (order Decapoda).

Genus Alphadrosrhavirus. The viruses in this genus have been detected in flies of various species (Diptera: Drosophilidae). The genome features an additional gene between the G and L genes in which there are two overlapping ORFs, each of which encodes a small hydrophobic protein with a strongly predicted transmembrane domain.

Genus Alphahymrhavirus. The viruses in this genus have been detected in ants and wasps (order Hymenoptera) and are distinct phylogenetically from rhabdoviruses assigned to the genus Betahymrhavirus.

Genus Betahymrhavirus. The viruses in this genus have been detected in wasps (order Hymenoptera) and are distinct phylogenetically from rhabdoviruses assigned to the genus Alphahymrhavirus. The genome features an additional gene between the M gene and G gene with two overlapping reading frames and a “slippery” sequence in the overlap region that could allow expression of the second ORF by ribosomal frame-shift.

Genus Betanemrhavirus. The viruses in this genus have been detected in parasitic roundworms (Ascaridida: Ascarididae). The genome features an additional gene between the P and M genes.

Genus Betapaprhavirus. The viruses in this genus have been detected in moths and butterflies (order Lepidoptera). The genome features an additional gene between the G and L genes encoding a small basic protein.

Genus Betaricinrhavirus. The viruses in this genus have been detected in hard ticks (family Ixodidae). The genome features an alternative ORF in the N gene, overlapping the end of the N ORF.