Viruses of this genus are distinguished from other hypoviruses based on polyprotein-derived phylogenetic trees and have a large genome size of over 15 kb.
Genome organization and replication
Monocistronic, positive-sense RNA genomes of 15.2 to 18.3 kb.
Species demarcation criteria
Members of different species are distinguished by the arbitrary threshold of less than 80% amino acid sequence identity in the whole RdRP-containing polyproteins.